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The exchange of gases within an animal burrow has been an extensive area of study due to the barrier that the burrowing medium (both aquatic and terrestrial) often imposes to the diffusion of gases.
Joseph M. Szewczak, Matching Gas Exchange in the Bat from Flight to Torpor, American Zoologist, Vol. 37, No. 1 (Feb., 1997), pp. 92-100 ...
1. Physiological methods applicable to scales between individual leaves and whole forests have the potential to improve substantially our understanding of ecosystem gas exchange. 2. We compared three ...